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Acknowledgments

This work was supported by the Imageomics Institute, which is funded by the US National Science Foundation's Harnessing the Data Revolution (HDR) program under Award #2118240 (Imageomics: A New Frontier of Biological Information Powered by Knowledge-Guided Machine Learning). This material is based in part upon work supported by the National Ecological Observatory Network (NEON), a program sponsored by the U.S. National Science Foundation (NSF) and operated under cooperative agreement by Battelle.

S. Record and A. East were additionally supported by the US National Science Foundation's Award No. 242918 and by Hatch project Award #MEO-022425 from the US Department of Agriculture’s National Institute of Food and Agriculture. M. Belitz was additionally supported by the US National Science Foundation's Award #2410152.

The Ohio Supercomputer Center was used for data storage. Ohio Supercomputer Center. 1987. Ohio Supercomputer Center. Columbus OH: Ohio Supercomputer Center. https://ror.org/01apna436.

We thank Hojun Song, Principal Investigator of the NEON Biorepository and Director of the ASU Biocollections, Isa Betancourt, Kelsey Yule, Chandra Earl, and all of the NEON Biorepository staff and collection managers for their stewardship of the collection on which this work depends, and for hosting us throughout this imaging effort. We are aslo grateful to the NEON field staff and parataxonomists who collect, sort, and pin specimens, and to the expert taxonomists who provide identifications. We also thank the NEON site hosts whose stewardship enables continental-scale sampling, Laura Nagel for overseeing NEON's ground beetle field protocols, and Paula Mabee and Kate Thibault for their input, support, and institutional leadership.

Any opinions, findings and conclusions or recommendations expressed in this material are those of the author(s) and do not necessarily reflect the views of the National Science Foundation or US Department of Agriculture.